Чекліст флори України. 5: родина Caryophyllaceae s. l. (incl. Illecebraceae) (Caryophyllales, Angiosperms)
DOI:
https://doi.org/10.32999/ksu1990-553X/2023-19-1-1Ключові слова:
анотований список, поширення, види, підвиди, рід, родина, систематика, номенклатура, синоніми, гербарні зразки, Dianthus, Cerastium, Otites, Silene, Gypsophila, Minuartia.Анотація
За сучасними оцінками, родина Caryophyllaceae налічує близько 90 родів та від 2200 до 3000 видів, поширених на всіх континентах, але переважно поза тропіками. Для флори України наводиться 49 родів (з них три потребують підтвердження) та 231 вид (з підвидами). Найчисельнішими на автохтонні види у флорі України є роди Dianthus – 49 видів (разом з підвидами), Cerastium – 23, Otites – 17, Silene – 17, Gypsophila – 11, Minuartia s. str. – 8 тощо. В той же час багато родів представлені одним або двома видами (Agrostemma, Bufonia, Carpophora, Cucubalus, Elisanthe, Heliosperma, Kohlrauschia, Lychnis, Myosoton, Paronychia, Pleconax та ін. В порівнянні з попереднім виданням чеклісту, відбулися суттєві зміни у видовому складі родів: Dianthus, Silene, Elisanthe, Minuartia, Stellaria, у багатьох з яких виділені нові роди: з роду Stellaria – Alsine, Hylebia, Rabelera; з Silene – Atocion; з Elisanthe – Carpophora, Silenanthe; з Minuartia – Cherleria, Sabulina). Суттєвих номенклатурних змін зазнали родові й видові назви та таксономічний статус багатьох видів. Багато видів, які раніше наводилися для України потребують підтвер-дження (наводяться лише за літературними даними, старими зборами, або на основі лише одного гербарного зразка): Dianthus corymbosus, D. giganteiformis, D. pinifolius, D. pontederae, D. seguieri, D. serotinus, D. spiculifolius, Eudianthe coeli-rosa, Pseudosaponaria pilosa (≡ Gypsophila pilosa), Kohlrauschia velutina, Melandrium astrachanicum, M. nemorale, Moehringia lateriflora, Polyschemone nivalis (≡ Silene nivalis), Sagina apetala, S. hawaiensis (= S. subulata), Silene thymifolia, Spergularia syvaschica, Viscaria alpina. Зниклим у флорі України є середньоєвропейський вид Dianthus grationopolitanus (= Dianthus caesius Sm.), який раніше наводився у всіх флористичних зведеннях для околиць м. Заліщики (с. Хрещатик Заставнівського району Чернівецької області). Вимерлим видом є Heliosperma arcanum (≡ Ixoca arcana), описаний Х. Запаловичем з околиць м. Заліщики (Тернопільська область), за зборами G. Zipser, які після 1855 року ніхто більше не знаходив.
Посилання
Andrienko, T.L., Blum, O.B., Vasser, S.P. (eds.). 1985. Priroda Ukrainskoi SSR. Rastitelnyi mir. Nakova dumka, 208 p. (in Russian)
Andrzhejowsky, A.L. (1869). Caryophyllaceae. In: Rogovych A.C. Obosrenie semennykh b vysshikh sporovykh rasteniy, vkhodjacshikh v sostav flory guberniy Kievskoho uchebnoho okruha: Volynskoi, Podolskoi, Kievskoi, Chernihovskoi i Poltavskoi. Kiev: izdatelstvo Kievskoho universiteta, 299 p. (In Russian)
APG IV (Angiosperm Phylogeny Group). (2016). An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants. Botanical Journal of the Linnean Society 181(1): 1–20. https://doi.org/10.1111/boj.12385
Aydin, Z., Ertekin, A. S., Långström, E. & Oxelman, B. (2014). A new section of Silene (Caryophyllaceae) including a new species from South Anatolia, Turkey. Phytotaxa 178: 98–112 https://doi.org/10.11646/phytotaxa.178.2.2
Bieberstein (Marschall), F.A. (1808). Flora taurico-caucasica, T. 1. Charkoviae: Typis Academicis, 428 p.
Bittrich, V. (1993). Introduction to Centrospermae. In: Kubitzki K., Rohwer J.G. and Bittrich V. [eds.]. The families and genera of vascular plants, vol. 2, Magnoliid, Hamamelid and Caryophyllid families. Berlin: Springer Verlag: 13, 19.
Chater, A.O. (1964). Lychnis L. In: Flora Europaea, vol. 1. Eds. Tutin T.G., Heywood V.H., Burges N.A., Valentine D.H., Walters S.M., Webb D.A. Cambridge: Cambridge University Press: 56–157.
Chater, A.O. & Walters, S.M. (1964a). Notes on European Silene. Feddes Repertorium 69: 46–49.
Chater, A.O. & Walters, S.M. (1964b). Silene L. In: Flora Europaea, vol. 1. Eds. Tutin T.G., Heywood V.H., Burges N.A., Valentine D.H., Walters S.M., Webb D.A. Cambridge: Cambridge University Press: 158–181.
Chopyk, V.I. (1969). Floristic zoning of the Ukrainian Carpatians. Ukrainian Botanical Journal 26(4): 3–15. (in Ukrainian)
Chowdhuri, P.K. (1957). Studies in the genus Silene. Notes from the Royal Botanic Garden, Edinburgh 22(3): 221–278.
Didukh, Ya.P. (2009). (ed.). Red data book of Ukraine. Plant kingdom. Kyiv: Globalkonsalting, 912 р. (in Ukrainian)
Dillenberger, M.S. & Kadereit, J.W. (2014). Maximum polyphyly: Multiple origins and delimitation with plesi-omorphic characters require a new circumscription of Minuartia (Caryophyllaceae). Taxon 63(1): 64–88. https://doi.org/10.12705/631.5
Eggens, F., Popp, M., Nepokroeff, M., Wagner, W.L. & Oxelman, B. (2007). The origin and number of intro-ductions of the Hawaiian endemic Silene species (Caryophyllaceae). American Journal of Botany 94: 210–218.
Erixon, P. & Oxelman, B. (2008). Reticulate or treelike chloroplast DNA evolution in Sileneae (Caryophyllaceae)? Molecular Phylogenetics and Evolution 48: 313–325.
Fateryga, A.V., Svirin, S.A., Evseenkov, P.E. & Yena, A.V. (2017). On the presence of Holosteum marginatum C.A.Mey. in the Crimea. Turczaninowia 20(2): 23–30. (in Russian) https://doi.org/10.14258/turczaninowia.20.2.3
Fedoronchuk, M.M. (1997a). Silene L. sensu lato in Ukraine: a taxonomic review of segregate genera Oberna Adans. and Pleconax Raf. (Caryophyllaceae). Ukrainian Botanical Journal 54 (1): 70–76. (in Ukrainian)
Fedoronchuk, M.M. (1997b). Silene L. sensu lato in Ukraine: a taxonomic review of genus Silene sensu stricto (Caryophyllaceae). Ukrainian Botanical Journal 54 (6): 557–564. (in Ukrainian)
Fedoronchuk, M.M. (2004). The systeme Caryophyllaceae Juss. s. l. of Ukrainian flora and critical revision of some generas. Naukowyi wisnyk Chernivetskoho Universitetu: Zbirnyk naukovyh prats 223: 171–190. (in Ukrainian)
Fedoronchuk, M.M. (2022a). Ukrainian flora checklist. 1: family Lamiaceae (Lamiales, Angiosperms). Chornomorski Botanical Journal 18(1): 5–27. https://doi.org/10.32999/ksu1990-553X/2022-18-1-1 (in Ukrainian)
Fedoronchuk, M.M. (2022b). Ukrainian flora checklist. 2: family Fabaceae (Fabales, Angiosperms). Chornomorski Botanical Journal 18 (2): 97–138. https://doi.org/10.32999/ksu1990-553X/2022-18-2-1 (in Ukrainian) Fedoronchuk, M.M. (2022c). Ukrainian flora checklist. 3: family Apiaceae (= Umbelliferae) and Araliaceae (Apiales, Angiosperms Chornomorski Botanical Journal 18 (3): 203–221. https://doi.org/10.32999/ksu1990-553X/2022-18-3-1 (in Ukrainian) Fedoronchuk, M.M. (2022d). Ukrainian flora checklist. 4: family Rosaceae (Rosales, Angiosperms). Chornomorski botanical journal 18 (4): 305–349. https://doi.org/10.32999/ksu1990-553X/2022-18-4-1
Fedoronchuk, M.M. & Didukh, Ya.P. (2002). Ecoflora Ukrainy. Volume 3. Kyiv: Fitosociotsentr, 496 p. (in Ukrainian)
Fedoronchuk, M.M. & Mosyakin, S.L. (2016). The genus Minuartia s. l. (Caryophyllaceae) in the flora of East-ern Europe: an overview of nomenclatural changes in the light od new molecular phylogenetic data. Ukrainian Botanical Journal 73(2): 134–143. https://doi.org/10.15407/ukrbotj73.02.134 (in Ukrainian)
Fenzl, E. (1833). Versuch einer Darstellung der geographischen Verbreitung- und Verheilungs-Verhältnisse der natürlichen Familie der Alsineen in der Polarregion und eines Theils der gemässigten Zone der alten Welt. Wien, S. 1–70.
Fior, S. & Karis, P.O. (2007). Phylogeny, evolution and systematics of Moehringia (Caryophyllaceae) as inferred from molecular and morphological data: a case of homology reassessment. Cladistics 23: 362–372.
Fior, S., Karis, P.O., Casazza, G., Minuto, L., Sala, F. (2006). Molecular phylogeny of the Caryophyllaceae (Caryophyllales) inferred from chloroplast matK and nuclear rDNA ITS sequences. American Journal of Botany 93: 399–411.
Fodor, S.S. (1974). Flora Zakarpattya. Lviv: Vyscha shkola, vydavnytstvo pry Lvivskomu derzh. Universyteti, 208 p. (in Ukrainian)
Frajman, B. & Oxelman, B. (2007). Reticulate phylogenetics and phytogeographical structure of Heliosperma (Sileneae, Caryophyllaceae) inferred from chloroplast and nuclear DNA sequences. Molecular Phyloge-netics and Evolution 43: 140—155.
Frajman, B., Eggens, F. & Oxelman, B. (2009a). Hybrid origins and homoploid reticulate evolution within Heliosperma (Sileneae, Caryophyllaceae) — a multigene phylogenetic approach with relative dating. Systematic Biology 58: 328–345.
Frajman, B., Heidari, N. & Oxelman, B. (2009b). Phylogenetic relationships of Atocion and Viscaria (Sileneae, Caryophyllaceae) inferred from chloroplast, nuclear ribosomal, and low-copy gene DNA sequences. Taxon 58: 811–824.
Friedrich, H.C. (1979). Familie Illecebraceae, Familie Caryophyllaceae. In: Hegi G. [ed.] Illustrirte Flora von Mitteleuropa 3, Teil 2. Berlin-Hamburg: Verlag Paul Parey: 749–762, 763–946.
Gams, H. (1953). Beiträge zur Kenntnis der arktisch-alpinen Saginen. Phyton 5: 107–117.
Golubev, V.N. (1995). New addenda to the Crimean flora. Botanical Journal 80(11): 46–54. (in Russian)
Gorshkova, S.G. (1936). Lychnis L. In: Flora SSSR, vol. 6. Eds. V.L. Komarov, B.K. Schischkin. Mosqua, Leningrad: Editio Academiae Scientarum URSS: 692–699. (in Russian)
Greenberg, A.K. & Donoghue, M.J. (2011). Molecular systematics and character evolution in Caryophyllaceae. Taxon 60(6): 1637–1652.
Greuter, W. (1995). Silene (Caryophyllaceae) in Grecce: a subgeneric and sectional classification. Taxon 44(4): 543–581.
Harbaugh, D.T., Nepokroeff, M., Rabeler, R.K., McNeill, J., Zimmer, E.A. & Wagner, W.L. (2010). A new line-age-based tribal classification of the family Caryophyllaceae. International Journal of Plant Sciences 171(2): 185–198.
Hartman, R.L., Thieret, J.W., Rabeler, R.K. (2005). Paronychia Miller. In: Flora of North America north of Mexico. Magnoliophyta: Caryophyllidae, pt 2 / Ed. by FNA Editorial Committee. New York; Oxford: Oxford Univ. Press 5: 30–43.
Heluta, V.P. (1989). Flora gribov Ukrainy. Muchnistorosyanye griby. Kiev: Naukova dumka, 256 p. (in Russian)
Herbich, F. (1859). Flora von Bukovina. Leipzig: A.Edelmann, I–VI+460 S.
Hernandez-Ledesma, P., Walter, G., Berendsohn, W.G., Borsch, T., Mering, S. von, Akhani, H., Arias, S., Castaneda-Noa, I., Eggli, U., Eriksson, R., Flores-Olvera, H., Fuentes-Bazan, S., Kadereit, G., Klak, C., Korotkova, N., Nyffeler, R., Ocampo, G., Ochoterena, H., Oxelman, B., Rabeler, R.K., Sanchez, A., Schlumpberger, B.O. & Uotila, P.A. (2015).Taxonomic backbone for the global synthesis of species diversity in the angiosperm order Caryophyllales. Willdenowia, 45: 281–383. https://doi.org/10.3372/wi.45.45301
Hutchinson, J. (1926). The Families of Flowering Plants, vol. I: Dicotyledons. London: McMillan and Co, 328 p.
Ikonnikov, S. (1973). Notae de Caryophyllaceae, 1. Novitates systematicae plantarum vascularium 10: 136–142. (in Russian)
Ikonnikov, S. (1975). Notae de Caryophyllaceae, 3. Novitates systematicae plantarum vascularium 12: 196–201. (in Russian)
Ikonnikov, S. (1976). Notae de Caryophyllaceae, 3. Novitates systematicae plantarum vascularium13: 113–120. (in Russian)
Ikonnikov, S. (1977). Notae de Caryophyllaceae, 5. Novitates systematice plantarum vascularium 14: 74–79. (in Russian)
Ikonnikov, S. (1978). Notae de Caryophyllaceae, 6. Novitates systematicae plantarum vascularium 15: 74–79. (in Russian)
Ikonnikov, S. (1984). Notae de Caryophyllaceae, 7. Novitates systematicae plantarum vascularium 21: 61–67. (in Russian)
Ikonnikov, S. (1987). Notae de Caryophyllaceae, 8. Novitates systematicae plantarum vascularium 24: 79–84. (in Russian)
Ikonnikov, S.S. (2004a.) Gypsophila L. In: Flora Europae Orientalis, vol. 11. Red. N.N. Tzvelev. Moscua-Petropoli: Oficina Editoria KMK: 257–265. (in Russian)
Ikonnikov, S.S. (2004b). Pleconax Raf. In: Flora Europae Orientalis, vol. 11. Red. N.N. Tzvelev. Moscua-Petropoli: Oficina Editoria KMK, p. 256–257. (in Russian)
Ivan, D. (1972). Răspindirea speciei Silene thymifolia Sibth. & Sm. pe litoralui mării Negre. Acta Horti Botanici Bucurestiensis 1970– 1971 10: 515–518.
Jalas, J. & Suominen, J. 1983. (Eds.) Atlas florae Europaeae. Distribution of vascular plants in Europe. Caryo-phyllaceae (Alsinoideae and Paronychioideae). Volume 6. Helsinki, 176 p.
Jalas, J. & Suominen, J. (Eds.) 1986. Atlas florae Europaeae. Distribution of vascular plants in Europe. Caryo-phyllaceae (Silenoideae). Volume 7. Helsinki, 229 p.
Kleopow, Ju.D. (1932). Review of representatives of Dianthus of Ukraine and contiguous localities. Visnyk Kyivskoho Botanichnogo Sadu 14: 99–140. (in Ukrainian)
Kleopow, Ju.D. (1936). To systematization and geography of Caryophyllaceae of the USSR. Journal Institute of Botany URSR 9(17): 91–126. (in Ukrainian)
Kleopow, Ju.D. (1939). News of flora of Prysyvashshya. Journal Institute of Botany URSR 21–22(29–30): 243–252. (in Ukrainian)
Klokov, M.V. (1922). Note about Gypsophila stepposa sp. n. Journal of Russian Botany Society 1921 6: 139–142. (in Russian)
Klokov, M.V. (1923). Gypsophila stepposa Klokov sp. n. Botanical materials (Leningrad) 4(11–12): 95–96. (in Russian)
Klokov, M.V. (1924). Gypsophila stepposa Klokov sp. n. Ukrainian Botanical Journal 2: 44–45 (in Ukrainian)
Klokov, M.V. (1947). New materials are to cognition of the Ukrainian flora. 2. Critical revision of family of Caryophyllaceae: generas of Cerastium L., Minuartia L., Herniaria L. Botanical Journal AN URSR 4(1–2): 60–75. (in Ukrainian)
Klokov, M.V. (1948). New materials are to cognition of the Ukrainian flora. 3. New species from families Caryophylaceae, Polygonaceae and Cruciferae. Botanical Journal AN URSR 5(1): 20–31. (in Ukrainian)
Klokov, M.V. (1952). Caryophyllaceae Juss. In: Flora URSR, vol. 4. Ed. M.I. Kotov. Kyiv: Vydavnytstvo AN Ukrainskoi RSR: 421–649. (in Ukrainian)
Klokov, M.V. (1974). Status praesens investigationis Caryophyllacearum Ucrainicarum. Novitates systematicae plantarum vascularium et non vascularium: 7–67. (in Russian)
Knjasev, M.S. (2012). Dianthus L. In: Conspectus florae Europae Orientalis, vol. 1. Red. N.N. Tzvelev. Petropoli-Mosqua: Consociatio editionum scientificarum KMK: 255–265. (in Russian)
Kool, A. (2012). Desert Plants and Deserted Islands: Systematics and Ethnobotany in Caryophyllaceae. Acta Universitatis Upsaliensis, Digital Comprehensive Summaries of Uppsala Dissertations from the Faculty of Science and Technology 972: 52.
Kotov, M.I. (1960). New plant of flora of URSR – Gypsophila of scorzonerifolia Ser. Ukrainian Botanical Journal 17(4): 75–79. (in Ukainian)
Kozhanchikov, V.I. (1970). Morphological and geographical studies of seeds of representatives of the family Caryophyllaceae Juss. European part of the USSR: Abstract of diss. … cand. biol. sciences., 19 p. (in Russian)
Kozhanchikov, V.I. (1975). Variability of morphological traits of seeds of representatives of the family Caryo-phyllaceae. Questions of Modern Morphology of Seed Plants. L.: Nauka, p.108–128. (in Russian)
Krupkina, L.I. (2004). Vaccaria N.M.Wolf. In: Flora Europae Orientalis, vol. 11. Red. N.N. Tzvelev. Moscua-Petropoli: Oficina Editoria KMK: 270–271. (in Russian)
Krupkina, L.I. (2012). Vaccaria N.M.Wolf. In: Conspectus florae Europae Orientalis, T. 1. Red. N.N. Tzvelev. Petropoli-Mosqua: Consociatio editionum scientificarum KMK, p. 253–254. (in Russian)
Krytska, L.I., Fedoronchuk, M.M., Tsarenko, O.V. & Shevera, M.V. (1999). Typification of species of vascular plants described from Ukraine: famile Caryophyllaceae Juss. (subfamile Silenoideae A.Br.). Ukrainian Botanical Journal 56(4): 402–410. (in Ukrainian)
Kulczyński, S. (1921). Fam. Caryophyllaceae. In: Flora Polska, T. 2. Red. Szafer W. Warszawa-Krakow: 125–184.
Kurtto, A. (2001a). Lychnis L. In: Flora Nordica (Chenopodiaceae to Fumariaceae). Vol. 2. Ed. B. Jonsel. Stockholm: 177–180.
Kurtto, A. (2001b). Silene L. In: Flora Nordica (Chenopodiaceae to Fumariaceae). Vol. 2. Ed. B. Jonsel. Stockholm: 180–202.
Kurtto, A. (2001c). Spergularia (Pers.) J.Presl et C.Presl In: Flora Nordica (Chenopodiaceae to Fumariaceae). Vol. 2. Ed. B. Jonsel. Stockholm: 92–96.
Kuzmina, M.L. (2002a). The sections Macrolepides (F.N.Williams) Klok. and Barbulatum F.N.Williams of the genus Dianthus (Caryophyllaceae) in East Europe and the Caucasus. Komarovia 2: 29–54.
Kuzmina, M.L. (2002b). The sections Carthusiani and Armerium of the genus Dianthus (Caryophyllaceae) in East Europe and the Caucasus. Komarovia 3: 85–102.
Kuzmina, M.L. (2004). Dianthus L. In: Flora Europae Orientalis, T. 11. Red. N.N. Tzvelev. Moscua-Peropoli: Oficina Editoria KMK: 273–297. (in Russian)
Ledebour, C.F. (1842). Flora Rossica. Enumeratio Plantarum, vol. 1. Stuttgartiae: Sumtibus Librariae E. Schweizerbart, 787 S.
Lidén, M., Popp, M. & Oxelman, B. (2001). A revised generic classification of the tribe Sileneae (Caryo-phyllaceae). Nordic Journal of Botany 20: 513–518.
Linnaeus, C. (1753). Species Plantarum ex hibens plantas rite cognitas, ad genera relata cum differentiis speci-ficis, vol. 1. Holmiae (Stockhollm): Impensis Laurentii Salvii, 560 p.
López González, G. (2010). Sobre el género Spergula L. [incl. Spergularia (Pers.) Pers. ex J.Presl & C.Presl, nom. cons.] (Caryophyllaceae) y sus especies en la Península Ibérica e Islas Baleares. Lagascalia 30: 7–18.
Löve, A.D. & Löve, D. (1948). Chromosome numbers of Northern Plant Species. Reykjavik, 75 p.
Löve, A.D., Löve, D. (1975). Cytotaxonomical atlas of the artic flora. Vaduz, 598 p.
Madhani, H., Rabeler, R., Pirani, A., Oxelman, B., Heubl, G. & Zarre, S. (2018). Untangling phylogenetic patterns and taxonomic confusion in tribe Caryophylleae (Caryophyllaceae) with special focus on generic boundaries. Taxon 67(1): 83–112. https://doi.org/10.12705/671.6 Margittai, A. (1927). Újabb adatok Turoczvármegye fórájához. (Neuere Beiträge zur Flora des Turóczer Komi-tates). Magyar botanikai lapok 25: 219–226.
Mattfeld, F. (1921). Gen. Scleranthus. Berichte der Deutschen Botanischen Gesellschaft 39: 275.
Mattfeld, F. (1922). Geographisch-genetische Untersuchungen über die Gattung Minuartia (L.) Hier. Repertori-um Specierum Novarum Regni Vegetabilis 15:1–228.
Mayol, M.& Rossello, J.A. (2000). Interpopulation esterase variation in the genus Petrocoptis A.Braun (Caryo-phyllaceae). Israel Journal of Plant Sciences 48(2): 79–89.
McNeill, J. (1962a). Taxonomic studies in the Alsinoideae. I. Generic and infra-generic groups. Notes from the Royal Botanic Garden, Edinburgh 24: 79–153.
McNeill, J. (1962b). Taxonomic studies in the Alsinoideae. II. A revision of the species in the Orient. Notes from the Royal Botanic Garden, Edinburgh 24: 241–426.
McNeill, J. (1980). A new species of Arenaria subgenus Odontostemma from Tibet, with a review of the status of the genus Gooringia (Caryophyllaceae). Botanical Journal of the Linnean Society 80(4): 371–378.
McNeil, J. & Bassett, I.J. (1974). Pollen morphology and the infrageneric classification of Minuartia (Caryo-phyllaceae). Canadian Journal of Botany 52(6): 1225–1231.
Meusel, H. & Werner, K. (1979a). Lychnis L. In: Illustrirte Flora von Mitteleuropa, Aufl. 2, Bd 3, Teil 2. Ed. G. Hegi. Berlin; Hamburg: Verlag Paul Parey: 1153–1175.
Meusel, H. & Werner, K. (1979b). Silene L. In: Illustrirte Flora von Mitteleuropa, Aufl. 2, Bd 3, Teil 2. Ed. G. Hegi. Berlin; Hamburg: Verlag Paul Parey, S. 1043—1153.
Morton, J.K. (1972). On the occurrence of Stellaria pallida in North America. Bulletin of the Torrey Botanical Club 99: 95–97.
Morton, J.K. (2005). Stellaria Linnaeus. In: Flora of North America north of Mexico. Ed. by FNA Editorial Committee, vol. 5, Magnoliophyta: Caryophyllidae, part 2. – New York; Oxford: Oxford University Press: 96–114.
Mosyakin, S.L. & Fedoronchuk, M.M. (1999). Vascular plants of Ukraine. A nomenclatural checklist, Kiev, xxiii + 345 p. http://dx.doi.org/10.13140/2.1.2985.0409
Mosyakin, S.L. & Fedoronchuk, M.M. (2015). New combinations for East European species of Sabulina (Caryo-phyllaceae). Phytotaxa, 231(1): P. 95–98. http://dx.doi.org/10.11646/phytotaxa.231.1.10
Nepokroeff, M. (2001). Origin of the Hawaiian subfam. Alsinoideae and preliminary relationships in Caryophyl-laceae inferred from matK and trnL C-F sequence data. American Journal of Botany Abstracts. Columbus: 130.
Nepokroeff, M., Wagner, W.L., Weller, S.G., Soltis, P.S., Zimmer, E.A., Sakai, A.K. & Soltis, D.E. (2001a). Diversification in the endemic Hawaiian subfam. Alsinoideae (Caryophyllaceae): evidence from nrDNA ITS, ETS sequences and morphology. Botany Abstracts: 129–130.
Nepokroeff, M., Wagner, W.L., Zimmer, E.A., Weller, S.G., Sakai, A.K. & Rabeler, R.K. (2001b). Origin of the subfam. Alsinoideae and preliminary relationships in Caryophyllaceae inferred from matK and trnL C-F sequence data. Botany: 130.
Neumayer, H. (1923). Einige Fragen der speziellen Systematik, erläutert an einer Gruppe der Gattung Silene. Oesterreichische Botanische Zeitschrift 72: 276–287.
Orlov, O.O., Shynder, O.I., Vorobjov, E.O. & Gryb, O.V. (2022). New floristic finds in the Forest-Steppe part of Zhytomyr Region. Ukrainian Botanical Journal 79(1): 6–26 [In Ukrainian]. https://doi.org/10.15407/ukrbotj79.01.006
Oxelman, B., Ahlgren, B. & Thulin, M. (2002). Circumscription and phylogenetic relationships of Gymnocarpos (Caryophyllaceae-Paronychioideae). Edinburgh Journal of Botany 59: 221–237.
Oxelman, B. & Liden, M. (1995). Generic boundaries in the tribe Sileneae (Caryophyllaceae) as inferred from nuclear rDNA sequences. Taxon 44(4): 525–542.
Oxelman, B., Liden, M. & Berglund, D. (1997). Chloroplast rps16 intron phylogeny of the tribe Sileneae (Caryophyllaceae). Plant Systematics and Evolution 206(1–4): 393–410.
Oxelman, B., Liden, M. & Jonsell, B. (2001). Proposal to conserve the name Viscaria against Steris (Caryophyl-laceae, Sileneae). Taxon 50(1): 281–282.
Oxelman, B., Liden, M., Rabeler, R.K. & Popp, M. (2000). A revised generic classification of the tribe Sileneae (Caryophyllaceae). Nordic Journal of Botany 20(6): 743–748.
Oksijuk, P.F. (1948). Caryologichne vyvchennya flory URSR. Botanical Journal AN URSR 5(1): 11–19. (in Ukrainian)
Paczosky, J.K. (1910). Osnovnye cherty razvitiya flory Jugo-Zapasdnoy Rossii. Zapiski Novorosiskoho obshchestva estestvoispytateley 34: 1–430. (in Russian)
Pawłowski, B. & Walas, J. (1949). Les associations des plantes vаsculaires des Monts de Czywczyn. Bulletin of the Polish Academy of Sciences, Biology Science 1: 1–181.
Pax, F. & Hoffmann, K. (1934). Caryophyllaceae. In: Die natürlichen Pflanzenfamilien. 2. Aufl., B. 16 C. Eds. Engler A., Prantl K. Leipzig: 275–364.
Pirani, A., Zarre, S., Pfeil, B.E., Bertrand, Y., Assadi, M. & Oxelman, B. (2014). Molecular phylogeny of Acan-thophyllum (Caryophyllaceae, Caryophylleae), with emphasis on subgeneric classification. Taxon 63: 592–607. https://doi.org/10.12705/633.39
Popp, M. & Oxelman, B. (2004). Evolution of an RNA polymerase gene family in Sileneae (Caryophyllaceae) – incomplete concerted evolution and topological congruence among paralogues. Systematic Biology 53: 914–932.
Popp, M., Gizaw, A., Nemomissa, S., Suda, J. & Brochmann, C. (2008). Colonization and diversification in the African‘sky islands’ by Eurasian Lychnis L. (Caryophyllaceae). Journal of Biogeography 35: 1016–1029.
POWO 2022. Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Available at: http://wwwplantsoftheworldonline.org (Accessed 15 April 2022 and 28 June 2022).
Prodan, J. (1953). Genul 95. Melandrium Rohl. In: Flora Republicii Populare Romine. Edit. Acad. RPR: 186–198.
Prodan, J. & Buia, Al. (1961). Flora Mica Ilustrata a Republicii populare Romãne. Ministerul Agriculturii. Editura Agro-Silvica. Bucuresti, 676 p. Prokudin, Yu.N. (ed.). (1987). Identification key to higher plants of Ukraine. Kyiv: Naukova dumka, 548 p. (in Russian)
Rabeler, R.K. (2005). Myosoton Moench. In: Flora of North America north of Mexico. Magnoliophyta: Caryophyllidae, pt 2, vol. 5. Ed. by FNA Editorial Committee. New York; Oxford: Oxford Univ. Press: 95–96.
Rabeler, R.K. & Hartman, R.L. (2005). Caryophyllaceae. In: Flora of North America north of Mexico Magnoliophyta: Caryophyllidae, pt. 2, vol. 5. Ed. by FNA Editorial Committee. New York: Oxford Univ. Press: 3–8.
Rautenberg, A., Filatov, D., Svennblad, B., Heidari, N. & Oxelman, B. (2008). Conflicting phylogenetic signals in the SIX1/Y1 gene in Silene. BMC Evolutionary Biology, 8: 299. https://doi.org/10.1186/1471-2148-8-299
Rautenberg, A., Sloan, D.B., Aldén, V. & Oxelman, B. (2012). Phylogenetic Relationships of Silene multinervia and Silene section Conoimorpha (Caryophyllaceae). Systematic of Botany 37: 226–237. https://doi.org/10.1600/036364412X616792
Rohrbach, P. (1868). Monographie der Gattung Silene (Diss.). Leipzig, 249 S.
Sadeghian, S., Zarre, S., Rabeler, R.K. & Heubl, G. (2015). Molecular phylogeny of Arenaria (Caryophyllaceae: tribe Arenarieae) and its allies inferred from nuclear DNA ITS and plastid DNA rps16 sequences. Botanical Journal of the Linnean Society 178: 648–669. http://dx.doi.org/10.1111/boj.12293
Schischkin, B.K. (1936). Caryophyllaceae Juss. In: Flora SSSR, vol. 6. Eds V.L. Komarov, B.K. Schischkin. Mosqua-Leningrad: Editio Academiae Scientarum URSS: 386–900. (in Russian)
Schishkin, B.K. (1947). Caryophyllaceae. In: Flora Kryma, vol. 1, vyp. 1. Ed. E.V. Wulf. Mosqua; Leningrad: Selkhozgiz: 117–171. (in Russian)
Schmalhausen, I.F. (1897). Flora Srednei i Jushnoi Rossii, Kryma i Severnoho Kavkaza: Rukovodstvo dlay opregelenia semehhykh i sporovykh rasteniy 2. Kiev, 889 p. (in Russian)
Sharples, M.T. & Tripp, E.A. (2019). Phylogenetic relationships within and delimitation of the cosmopolitan flowering plant genus Stellaria L. (Caryophyllaceae): Core stars and fallen stars. Systematic Botany 44(4): 857–876 (14 November 2019). https://doi.org/10.1600/036364419X15710776741440
Sheen, A.-C., Brochman, C., Brysting, A.K., Elven, R., Morris, A., Soltis, D.E., Soltis, P.S. & Albert, V.A. (2004). Northern hemisphere biography of Cerastium (Caryophyllaceae): insights from phylogenetic analysis of noncoding plastid nucleotide sequences. American Journal of Botany 91(6): 943–952.
Smissen, R.D., Clement, J.C., Garnock-Jones, P.J. & Chambers, G.K. (2002). Subfamilial relationships within Caryophyllaceae as inferred from 5' ndhF sequences. American Journal of Botany 89(8): 1336–1341.
Smissen, R.D. & Garnock-Jones, P.J. (2002). Relationships, classification, and evolution of Scleranthus (Caryo-phyllaceae) as inferred from analysis of morphological characters. Botanical Journal of the Linnean Society 140(1): 15–29.
Sokolova, I. (2001). Quid est Cerastium villosum Stev. (Caryophyllaceae). Novitates systematicae plantarum vascularium 33: 87–89. (in Russian)
Sokolova, I.V. (2004). Cerastium L. In: Flora Europae Orientalis, vol. 11. Red. N.N. Tzvelev. Moscua-Petropoli: Oficina Editoria KMK: 157–171. (in Russian)
Sokolova, I.V. (2012). Cerastium L. In: Conspectus florae Europae Orientalis, T. 1. Red. N.N. Tzvelev. Petropoli-Mosqua: Consociatio editionum scientificarum KMK: 210—255. (in Russian)
Steven, C. (1856). Verzeichniss auf der Taurischen Halbinsel wildwachsenden Pflanzen. Bulletin de la Société impériale des naturalistes de Moscou 29(2): 234–334.
Stoiko, S. & Tasenkevitsch, L. (1991). Pflanzengeographische Stellung und Schulz von Flora und Vegetation der Ukrainischen Karpaten. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 128:165–177.
Takhtadjan, A. (1997). Diversity and classification of flowering plants. New-York: Columbia University Press, 643 p.
Tokhtar, V.K. (1996). Addition to the adventive flora of southeast of Ukraine. Ukrainia Botanical Journal 53(6): 687–690. (in Ukrainian)
Tsvelev, N.N. (1984). On two species new to the European part of the USSR from the genera Melandrium (Caryophyllaceae) and Veronica (Scrophulariaceae). Botanical Journal 69(9): 1255–1260. (in Russian)
Tsvelev, N.N. (2000). Notulae de generibus nonnullis familiae Сaryophyllaceae sensu lato in Europa Orientali. Novitates Systematicae Plantarum Vascularium 32: 26–36. (in Russian)
Tsvelev, N.N. (2001). De generibus tribus Sileneae DC. (Caryophyllaceae) in Europa Orientali. Novitates Systematicae Plantarum Vascularium 32 90–113. (in Russian)
Tsvelev, N.N. (2002). About some generas of family Caryophyllaceae in Eastern Europe. Botanical Journal 87(3): 120–130. (in Russian)
Tsvelev, N.N. (2004a). Carpophora Klotzsch. In: Flora Europae Orientalis, vol. 11. Red. N.N. Tzvelev. Moscua-Petropoli: Oficina Editoria KMK: 224–225. (in Russian)
Tsvelev, N.N. (2004b). Corrigiola L. In: Flora Europae Orientalis, vol. 11. Red. N.N. Tzvelev. Moscua-Petropoli: Oficina Editoria KMK: 132–133. (in Russian)
Tsvelev, N.N. (2004с). Elisanthe (Fenzl) Fenzl. In: Flora Europae Orientalis, vol. 11. Red. N.N. Tzvelev. Moscua-Petropoli: Oficina Editoria KMK: 224. (in Russian)
Tsvelev, N.N. (2004d). Hylebia (Koch) Fourr. In: Flora Europae Orientalis, vol. 11. Red. N.N. Tzvelev. Moscua-Petropoli: Oficina Editoria KMK: 153–154. (in Russian)
Tsvelev, N.N. (2004e). Illecebraceae R.Br., nom. conserv. In: Flora Europae Orientalis, vol. 11. Red. N.N. Tzvelev. Moscua-Petropoli: Oficina Editoria KMK: 124–138. (in Russian)
Tsvelev, N.N. (2004f). Ixoca Raf. In: Flora Europae Orientalis, vol. 11. Red. N.N. Tzvelev. Moscua-Petropoli: Oficina Editoria KMK: 226–227. (in Russian)
Tsvelev, N.N. (2004g). Minuartia L. In: Flora Europae Orientalis, vol. 11. Red. N.N. Tzvelev. Moscua-Petropoli: Oficina Editoria KMK: 191–203. (in Russian)
Tsvelev, N.N. (2004h). Oberna Adans. In: Flora Europae Orientalis, vol. 11. Red. N.N. Tzvelev. Moscua-Petropoli: Oficina Editoria KMK: 229–233. (in Russian)
Tsvelev, N.N. (2004i). Otites Adans. In: Flora Europae Orientalis, vol. 11. Red. N.N. Tzvelev. Moscua-Petropoli: Oficina Editoria KMK: 247—256. (in Russian)
Tsvelev, N.N. (2004j). Sagina L. In: Flora Europae Orientalis, vol. 11. Red. N.N. Tzvelev. Moscua-Petropoli: Oficina Editoria KMK, p. 186–191. (in Russian)
Tsvelev, N.N. (2004k). Scleranthus L. In: Flora Europae Orientalis, vol. 11. Red. N.N. Tzvelev. Moscua-Petropoli: Oficina Editoria KMK: 203–206. (in Russian)
Tsvelev, N.N. (2004l). Spergularia (Pers.) J. & C.Presl. In: Flora Europae Orientalis, vol. 11. Red. N.N. Tzvelev. Moscua-Petropoli: Oficina Editoria KMK: 127–132. (in Russian)
Tsvelev, N.N. (2012a). Oberna Adans. In: Conspectus florae Europae Orientalis, т. 1. Red. N.N. Tzvelev. Petropoli-Mosqua: Consociatio editionum scientificarum KMK: 237–239. (in Russian)
Tsvelev, N.N. (2012b). Stellaria L. In: Conspectus florae Europae Orientalis, т. 1. Red. N.N. Tzvelev. Petropoli-Mosqua: Consociatio editionum scientificarum KMK: 205–208. (in Russian) Tutin, T.G. (1964). Dianthus L. In: Flora Europae, vol. 1. Eds. Tutin, T.G., Heywood, V.H., Burges, N.A., Moore, D.M., Valentine, D.H., Walters, S.M., Webb, D. Cambridge: Univer. Press: 188–204.
Tutin, T.G., Heywood, V.H., Burges, N.A., Moore, D.M., Valentine, D.H., Walters, S.M. & Webb, D. (eds.). (1964). Flora Europaeae, vol. 1 (Lycopodiaceae to Platanaceae). Cambridge: University Press, 455 p.
Vitova, J., Vit, P. & Suda, J. (2015). Rare occurrence of reciprocal hybridization in a sympatric population of the Czech stenoendemic Dianthus arenarius subsp. bohemicus and widespread D. carthusianorum. Preslia 87: 329–345.
Whitehead, F.H. & Sinha, R.P. (1967). Taxonomy and taximetricks of Stellaria media (Linnaeus) Vill., S. neglecta Weihe and S. pallida (Dumort.) Piré. New Phytology 66:769–784.
Williams, F.N. (1893). The desintegration of Lychnis. London Journal of Botany 31: 167–171.
Williams, F.N. (1898). A revision of the genus Arenaria. Botanical Journal of the Linnean Society 33: 326–437.
Zapałowicz, H. (1911). Conscpectus florae Galiciae criticus. Krytyczny przeglad roślinnośti Galicyi, T. 3. Krakowie: Nakladen Akademii Umiejętności, 252 s. Ziman, S.N. (1987).
Caryophyllaceae. In: Prokudin Yu.N. (Ed.) 1987. Identification key to higher plants of Ukraine. Kyiv: Naukova dumka: 65–83. (in Russian)
